My step dad is a wedding photographer but his true passion is nature photography so every second he gets he is out taking photos. What are some types of places that he could sell them to?

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you can check out: http://www.fineartamerica.com/ or you can sell them at local flee markets.

He could use a site like istockphoto and sell his pictures as stock. Basically someone pays a fee to be allowed to use and download his picture. People who have quality images can make a lot of money this way. Otherwise he should invest in getting some quality prints made and sell them himself either on a personal website or an online artist's market like Etsy.